Twitter, ahead of its eagerly-anticipated listing, revealed greater reliance on mobile advertising in an amended IPO filing – but not enough to stop Q3 losses tripling year-on-year.

More than 70 per cent of the company’s advertising revenue came from phones and tablets during the three months ended September, up from 65 per cent the previous quarter.

And although Q3 revenue more than doubled year-on-year, to $168.6 million, net losses jumped from $21.6 million to $64.6 million over the same period.

Twitter attributes the widening losses to increased spending on sales and marketing, which shot up from $23.7 million to $61.2 million.

Twitter reports the number of people using its service is more than 230 million people per month, up from about 218 million when the company disclosed its IPO filing on 3 October.

“Near-term you’re going to see this company grow very quickly,” Robert Peck, an analyst at SunTrust Robinson Humphrey, told Bloomberg in an interview before the amended filing was released. “Before they can be profitable they still have a long way to invest in their infrastructure and the future of the platform.”

The amended filing also revealed that JPMorgan Chase’s alternative asset management arm holds 10.3 per cent in Twitter.

Other shareholders include co-founder Evan Williams (12 per cent) and Rizvi Traverse (17.6 per cent), which is run by Hollywood and Silicon Valley financier Suhail Rizv.

CEO Dick Costolo, an early angel investor, owns 1.6 per cent.

Other major stakeholders include Spark Capital (6.8 per cent) and Benchmark Capital (6.6 per cent). Union Square Ventures owns 5.9 per cent.

In a blow to Nasdaq, which bungled Facebook’s listing in 2012, Twitter will trade on the New York Stock Exchange.

In its earlier filing, Twitter said it planned to raise $1 billion from going public (despite the recurring losses).

According to Bloomberg sources, Twitter is expected to kick off its investor roadshow on 28 October followed by a mid-November IPO.